EQUUS

Seeing spots

For as long as people have depicted horses in art, we’ve been showing them with spots.

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For as long as people have depicted horses in art, we’ve been showing them with spots.

Beginning years near what ago, about is people now 29,000 Pech living Merle, France, walked deep into a cave and painted their world on the walls. Together with numerous handprints, they created more than 800 distinct images, including mammoths, bison, aurochs, a bear---and horses drawn with dark outlines filled in with numerous round black spots, similar to the leopard-spotted horses we see today.

For a long time, scholars speculated about the spotted horses of Pech Merle. Ice Age humans painted many horses---1,250 have been documented in caves from Spain into eastern Russia ---but all others where coat colors can be identified are black, brown, bay, grullo or dun. Indeed, in this time before domesticat­ion, it was believed, these were the only equine coat colors that existed. With the assumption that the Pech Merle artists could not have seen real leopard-spotted horses, explanatio­ns for their paintings ranged from hallucinat­ions to artistic license.

In 2011, however, Stanford University researcher­s offered another possibilit­y: The artists may have been simply painting what they saw. Their genetic analysis showed that four of 10 horses they sampled from Pleistocen­e Europe carried the LP gene, while none of the six Siberian samples from the same period had it. LP is the leopard complex gene, which can produce leopard spotting, white blankets with spotting, and other color patterns we now associate mainly with Appaloosas and a few other breeds.

Headlines in the popular press in November 2011 made declaratio­ns like, “Spotted Horses in Cave Art Weren’t Just a Figment, DNA Shows” (The New York Times) and “Cave Paintings

Showed True Colors of Stone Age Horses” (Science Now). But did they?

Not necessaril­y, says Phillip Sponenberg, DVM, PhD, of the Virginia–Maryland Regional College of Veterinary Medicine and author of Equine Color Genetics. “Even though LP existed, the other question is how fully it was expressed,” he says. “Many LP horses only minimally express spotting, so basically they are not all that spotted. Further research might shed light on the modifiers that help to more fully express the spotting.”

French archaeolog­ist Jean Clottes noted that the Pech Merle artists painted spots both inside and outside the outlines of their horses, “giving them a meaning or role other than

purely descriptiv­e,” he told a reporter for the Los Angeles Times.

We may never understand the inspiratio­n behind the spotted horses of Pech Merle. “It depends on what was going through the artist’s mind, and that’s tough to figure out,” says Sponenberg. “So, while it could be that such spotted horses were out there, it equally could be artistic license.”

One thing is certain: For as long as people have known horses, we’ve been drawing and painting them with spots.

Reference: “Genotypes of predomesti­c horses match phenotypes painted in Paleolithi­c works of cave art,” Proceeding­s of the National Academy of Sciences, November 2011

 ??  ?? About 29,000 years ago: Paintings of spotted horses and human hands at the Pech Merle cave in France
About 29,000 years ago: Paintings of spotted horses and human hands at the Pech Merle cave in France
 ??  ?? 1047: From “Les quatre Cavaliers. Apoc. VI” (“The Four Horsemen of the Apocalypse VI”), an illustrati­on from the book The Beatus of Facundus Artist: Facundus, for Ferdinand I and Queen Sancha
1047: From “Les quatre Cavaliers. Apoc. VI” (“The Four Horsemen of the Apocalypse VI”), an illustrati­on from the book The Beatus of Facundus Artist: Facundus, for Ferdinand I and Queen Sancha
 ??  ?? 1650: “Landscape With Two Horses,” Dutch, oil on panel Artist: Nicolaes Pieterszoo­n Berchem
1650: “Landscape With Two Horses,” Dutch, oil on panel Artist: Nicolaes Pieterszoo­n Berchem
 ??  ?? 1296: “Grooms and Horses,” from the Yuan Dynasty in China; ink and color on paper Artist: Zhao Mengfu
1296: “Grooms and Horses,” from the Yuan Dynasty in China; ink and color on paper Artist: Zhao Mengfu
 ??  ?? About 1650 to 1654: “The ‘Piebald’ Horse,” Dutch, oil on canvas Artist: Paulus Potter
About 1650 to 1654: “The ‘Piebald’ Horse,” Dutch, oil on canvas Artist: Paulus Potter
 ??  ?? Circa 1750: “Scipio, a spotted hunter, the property of Colonel Roche,” English, oil on canvas Artist: Thomas Spencer
Circa 1750: “Scipio, a spotted hunter, the property of Colonel Roche,” English, oil on canvas Artist: Thomas Spencer
 ??  ?? Circa 1882 to 1892: “Equestrien­ne ( At the Cirque Fernando),” French, oil on canvas Artist: Henri de Toulouse-Lautrec
Circa 1882 to 1892: “Equestrien­ne ( At the Cirque Fernando),” French, oil on canvas Artist: Henri de Toulouse-Lautrec
 ??  ?? 1817: “Two Riders in a Landscape,” Dutch, oil on panel Artist: Anthony Oberman
1817: “Two Riders in a Landscape,” Dutch, oil on panel Artist: Anthony Oberman
 ??  ?? 1653: “The Spotted Horse,” Dutch, oil on panel Artist: Paulus Potter
1653: “The Spotted Horse,” Dutch, oil on panel Artist: Paulus Potter
 ??  ?? 1912: “Jumping Horse,” German, oil on canvas Artist: Franz Marc
1912: “Jumping Horse,” German, oil on canvas Artist: Franz Marc
 ??  ?? Circa 1850 to 1900: “Spotted Horse,” possibly from Pennsylvan­ia, in pen and ink plus watercolor on paper
Circa 1850 to 1900: “Spotted Horse,” possibly from Pennsylvan­ia, in pen and ink plus watercolor on paper
 ??  ?? 1876: From “Sketchbook,” Kiowa, graphite and colored pencil on paper Artist: Koba
1876: From “Sketchbook,” Kiowa, graphite and colored pencil on paper Artist: Koba
 ??  ?? 1984: Untitled, from the Lascaux Series Artist: Elaine de Kooning, color lithograph on wove paper, photo by Laura Shea
1984: Untitled, from the Lascaux Series Artist: Elaine de Kooning, color lithograph on wove paper, photo by Laura Shea

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