Ledecky takes silver in 400 free

American and defending Olympic champion Katie Ledecky was shocked in the women’s 400-meter freestyle in the Tokyo Games on Monday. She finished second to Australia’s Ariarne Titmus.

Titmus, who trailed Ledecky at the halfway mark of the race, turned on the speed to finish with a time of 3 minutes, 56.69 seconds, good for second-fastest all-time. Ledecky remains the world-record holder in the event.

Ledecky finished with the fourthfastest time ever, 3 minutes, 57.36 seconds.

“I fought tooth and nail,’’ Ledecky said. “She definitely swam a really smart race. She was really controlled up front. I felt pretty smooth and strong going out and flipped at the 300 and it was like, ‘Oh, she’s right there.’”

However, Team USA took gold in the men’s 4x100-meter freestyle relay.